Beyond the styling, the updates that matter are in the drivetrain, the suspension geometry and the towing package.
The headline is the twin-turbo V6, but the more interesting change is the electronic disconnecting sway bar, which lets the car stay flat on the highway and still articulate off it.
For the buyer
If your Land Cruiser spends most of its life on tarmac, the GX-R specification is the sweet spot. Higher trims add comfort you will notice and off-road hardware you may not use.